Douglas Carswell has become UKIP's first elected MP by claiming almost 60% of the vote in the Clacton by-election.
The former Conservative MP - who triggered the vote after defecting to UKIP - won 21,113 votes, compared to 8,709 for Conservative candidate Giles Watling.
In claiming victory in the landslide poll, Mr Carswell promised voters: "I will not let you down". He added that UKIP "must be a party for all Britain, and all Britons".
UKIP leader Nigel Farage said: "Congratulations Douglas Carswell, a brave and honourable man who has a just reward."
The result capped a dramatic few hours for UKIP, which pushed Labour right to the edge of defeat in the by-election at Heywood and Middleton.
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Gallery: Highs And Lows Of The By-Elections
Douglas Carswell has said "there is nothing that we cannot achieve" after winning the Clacton by-election with a staggering 12,404 majority.
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UKIP's first elected MP to Parliament forced the election after defecting from the Tories.
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Mr Carswell was the favourite to win, but there were still nerves on the night.
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Mr Carswell took almost 60% of the vote.
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Party leader Nigel Farage said: "Congratulations Douglas Carswell, a brave and honourable man who has a just reward."
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In the Heywood and Middleton by-election triggered by the death of Labour MP Jim Dobbin, Liz McInnes retained the seat for Labour in a closely fought contest.
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Ms McInnes won with 11,633 votes, defeating UKIP's John Bickley on 11,016. Ed Miliband's party's majority had been almost 6,000 in 2010, but a UKIP surge saw a 17.65% swing to Mr Farage's party.
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Despite UKIP's loss in the Lancashire constituency, Mr Bickley told Sky News "the Labour vote had collapsed".
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Mr Farage went on to tell Sky News: "We are ripping lumps out of the old Labour vote in the north of England. The truth of what has happened in the North today is that if you are anywhere north of Birmingham, if you vote Conservative you get Labour."
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He added: "And the reason we haven't won up there, despite a fantastic campaign, is that too many people have stuck with the Conservatives, not recognising that UKIP is now the challenger to Labour in every urban seat in the north of England."
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Turnout in Clacton was 51.2%, while in Heywood and Middleton it was just 36%.
Turnout was lower than expected in Greater Manchester, where just 36% of voters went to the polls.
Labour's Liz McInnes gained 11,633 votes in Heywood and Middleton, with UKIP's John Bickley finishing second with 11,016 votes.
The result was far closer than anticipated. UKIP requested a recount before Labour was confirmed as the winner.
The Conservatives scored 3,496 votes and the Liberal Democrats 1,457.
In her victory speech, Ms McInnes claimed the by-election result was a win for the NHS.
"The people gave their backing to Ed Miliband's plans for an NHS with the time to care," she said.
"They have rejected a Tory government that is only standing up for the privileged few."
But Sky's Jon Craig said the dramatic decline of Labour's majority in Heywood would trigger renewed debate about Mr Miliband's leadership.
"UKIP have done better here than even they thought they would," he said.
"They've come within a whisker of pulling off a sensational by-election victory."
The polls closed in Heywood and Middleton, and in Clacton, at 10pm on Thursday.
Voters in Heywood cast their ballot for a new MP following the death of Labour's Jim Dobbin last month.
Mr Dobbin held the seat from 1997 and was returned in 2010 with a significant majority.
The vote in Clacton was triggered in August by Mr Carswell's defection to UKIP.
Mr Carswell won the seat in 2010 with a majority of more than 12,000, having previously served as Conservative MP for Harwich until the constituency was abolished by boundary changes.
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Gallery: The UKIP History In Pictures
1993: UKIP is founded by Alan Sked in response to the Maastricht Treaty, which set out the modern day EU and paved the way for the Euro. He left the party in 1997 saying it had become a "racist party for the far-right". He is now the leader of New Deal, which has been called UKIP of the Left.
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1999: The party takes its first three seats in the European Parliament, under the leadership of the millionaire businessman Michael Holmes. Nigel Farage is one of those MEPs.
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2000: Michael Holmes resigns and Jeffrey Titford takes over as leader of UKIP. He leads the party to field 420 candidates at the 2001 General Election and secure 1.5% of the vote.
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2002: Former Conservative Roger Knapman takes over at the helm.
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2004: The party wins 12 seats at the European Elections, among the UKIP MEPs is the chat show host Robert Kilroy Silk.
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2005: Growing speculation Robert Kilroy Silk will take on the leadership comes to nothing and he announces he is leaving the party, calling it a "joke", setting up his own party, Veritas.
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2006: In a radio interview David Cameron calls UKIP members "fruitcakes, loonies and closet racists, mostly". It's the same year Nigel Farage is elected leader with 45% of the votes. Mr Farage drives an armoured vehicle to the Conservative Party Conference demanding an apology.
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2009: UKIP wins 13 seats at the European Elections but Mr Farage steps down as leader so he can concentrate on preparing for the General Election.
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2010: Nigel Farage decides to oppose House of Commons Speaker John Bercow in Buckingham - not the done thing. He fails to win the seat and goes on to reject the party's manifesto as "486 pages of drivel".
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2011: Ed Miliband hits the campaign trail at the Barnsley by-election (pictured) but UKIP candidate comes second to Labour, indicating the party presents a challenge to both Left and Right.
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February 2013: Diane James wins UKIP's highest by-election showing with 27.8% of the vote at Eastleigh. The Liberal Democrats hold the seat.
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September 2013: MEP Godfrey Bloom quits the party after provoking a row when he called women party activists, who didn't clean behind their fridges, "sluts". It came shortly after he made a reference to "bongo-bongo land".
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May 2014: UKIP becomes the first party other than Labour or the Conservatives in more than a century to win the majority share of the vote in a UK election at the local and European elections. Mr Farage claims he delivered the "earthquake" he promised.
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August 2014: Conservative MP Douglas Carswell announces he is defecting to UKIP triggering a by-election in Clacton.
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September 2014: Conservative MP Mark Reckless follows Mr Carswell and defects to UKIP on the eve of the Tory party conference, triggering a by-election in Rochester and Strood.
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October 2014: Nigel Farage announces he has parked his tanks on Labour's lawns as he joins the campaign trail in Heywood and Middleton, where a by-election is triggered by the death of the Labour MP Jim Dobbin, on the same day as the Clacton vote.
As he made his way to the Clacton count, Mr Carswell said he still had some "good friends" on the Tory benches, adding "friendship means a lot to me".
Asked by Sky News whether he could look them in the eye, he said: "I can certainly look myself in the mirror every morning."
He dismissed Tory claims that votes for UKIP would be a boost to Mr Miliband, claiming that he had "good data" that around half of his supporters had previously backed Labour.
More follows...
:: Watch full coverage of both by-elections in a special Sky News programme through the night - available on skynews.com, Sky News for iPad and on Sky 501, Virgin Media 602, Freesat 202 and Freeview 132.
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Gallery: UKIP Leader Nigel Farage: A Profile
Nigel Farage has been married twice and has four children, two sons from his first marriage and two daughters from his second. He continues to live in Kent and cites fishing, country sports, traditional English pubs and getting Britain out of the EU as his interests.
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MP Farage was born on April 3, 1964, in Herne, Kent, to stockbroker Guy Oscar Justus Farage and Barbara Stevens. His father was, reportedly, an alcoholic and left the family home when Mr Farage was just five years old. (Pic: Wikimedia Commons)
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Despite professing himself the head of the "people's army" and enemy of the "establishment", the young Nigel Farage was actually educated at public school Dulwich College.
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He decided not to go to university but opted instead for a career in the City as a commodities trader where he worked for two decades.
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Mr Farage started out as a Conservative but left the party in 1992 in protest at John Major's signing of the Maastricht Treaty. In 1993 he became a founder member of UKIP with the sole aim of getting Britain out of the European Union. Since taking over as leader in 2006 he has tried to claim the party as the only "real voice of opposition". There is, he says, "a cigarette paper between" the Labour, Conservative and Liberal Democrat parties.
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He was elected to the European Parliament as MEP for the South East of England in 1999 and held on to his seat in the elections of 2004 and 2009. In 2009 it was reported he claimed he had received a total of £2m of taxpayers money for staff, travel and other expenses. He has subsequently denied this but in an interview with Sky's Dermot Murnaghan in April 2014 said he was given £3,580 a month like all MEPs to spend how he wished.
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The UKIP leader has cemented an image of a man of the people and has been described as the politician with whom most people would like to share a pint. This is possibly because wherever he goes he is, in fact, pictured having a pint.
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... here he is pictured toasting with a landlord in the pub the night before ruling out running for the Newark by-election in April 2014.
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And here he is sharing a pint in February 2014 with volunteers helping during the flooding on the Somerset Levels.
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He also likes a cigarette.
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Mr Farage's campaign trail hat of choice is the Fedora, however, he is often seen out and about in a flat cap. He has run unsuccessfully for a seat in parliament a number of times, however, did deliver a resounding victory in the 2009 European elections capturing 16.5% of the vote - second only to the Tories.
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His most recent attempt at a seat in the Commons came at the 2010 General Election when he challenged the Tory MP and House of Commons Speaker John Bercow in Buckingham. He came third. In the same year he released his memoirs of founding UKIP and his fight against "establishment" political parties called Fighting Bull. He ruled out running for a seat in the Newark by-election on April 30, 2014.
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On May 6, 2010 - the morning of the General Election - Mr Farage was injured when a two-seater plane he was in crashed. He broke his sternum and ribs, and punctured his lung. An inquiry found that a banner the plane had been towing got caught in the tailplane, causing it to ditch. It was his second health scare - he was diagnosed with testicular cancer in his 20s but made a full recovery.
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Mr Farage has never been short of a headline - largely owing to a tasty choice of phrase and his strong views. In April 2014 he was sharply criticised for naming Vladmir Putin as the leader he admired the most and for praising the Russian President for being "brilliant" in his political manoeuvering over the Syrian crisis.
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He and his members have continually courted controversy, with one candidate telling the actor and comedian Lenny Henry to go and live in a "black country" and a councillor saying flooding was God's payback for David Cameron's gay marriage legislation. Mr Farage has also been criticised for employing his second wife, German-born Kirsten Mehr, as his European Parliamentary Secretary. He has said no one else would want the job. He was also branded "racist" for a UKIP European election campaign poster unveiled in April 2014.
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His views have made him unpopular with some. In May 2013 the UKIP leader had to be rescued by police in Edinburgh after being mobbed by protesters after a news conference ahead of an Aberdeen by-election.
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However, in the run-up to the May 2014 local and European elections Mr Farage's popularity surged. He triumphed in two televised debates with the Liberal Democrat leader Nick Clegg on Britain's membership of the EU.
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Mr Farage led the party to victory in the May vote. UKIP became the first party other than the Tories or Labour in more than a century to top a UK election with a 27.49% share of the vote. Significantly the party performed well in northern Labour heartlands.
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By September Mr Farage had pulled off a significant coup by securing the defection of two Conservative MPs, Douglas Carswell and Mark Reckless, and a £1m donation from the former Tory donor Arron Banks.
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